Contributions to Achenium Leach, 1819 (Coleoptera, Staphylinidae, Paederinae) fauna of Turkey

In this study, total of 388 specimens of the genus Achenium Leach, 1819 (Coleoptera: Staphylinidae) were collected by aspirator and sifter methods from different provinces of Aegean and Central Anatolia Regions of Turkey between 2014 and 2019. As a result of the field studies, five species of the genus were recorded. These species wereAchenium anatolicum Jarrige, 1952; A. humile (Nicolai, 1822); A. propontiacum Bordoni, 2009; A. scimbalioides Koch, 1937 and A. turcicum Coiffait, 1971. Of the recorded species; A. humile from the Aegean Region and A. turcicum from Central Anatolia Region were the first recorded for the area. A. turcicum was reported for the first time outside the type locality and its distribution was found to be much wider than previously known. A. turcicum and A. anatolicum were the endemic species for Anatolia. Research Article Article History Received : 11.02.2020 Accepted : 08.05.2020


INTRODUCTION
According to a recent comprehensive revision by , the genus Achenium Leach, 1819 (Staphylinidae, Paederinae) contains 53 species worldwide. In the meantime, a new species were described from Russia (Solodovnikov & Li, 2013). The biodiversity center of Achenium is the Mediterranean countries. Up to date, 11 species of the genus Achenium were recorded from Turkey, two of them are known from only this country Anlaş et al., 2011;Schülke & Smetana, 2015).
Despite many recent contributions, the current knowledge of the Achenium fauna of Turkey is far from complete. The main purpose of this study was to make a contribution to fauna of Achenium of Turkey.

MATERIAL and METHODS
The material has been obtained by aspirator and sifter methods in different field studies between 2014-2019 in Aegean and Central Anatolia Regions. Specimens were examined using a Stemi 508 microscope (Zeiss, Oberkochen, Germany). Photographs of the habitus, forebody and aedeagus were taken with a digital camera (Zeiss Axiocam ERC5s) All photographs were edited with the Helicon Focus v. 6, and Corel Draw v. X5 software.

RESULTS
In this study, five species of the genus Achenium belonging to the subfamily Paederinae were reported from Aegean and Central Anatolia regions of Turkey.
Amongst them, A. humile from the Aegean Region and A. turcicum from Central Anatolia Region were recorded for the first time. A. turcicum was reported for the first time outside the type locality and its distribution was found to be much wider than previously known. Achenium anatolicum Jarrige, 1952 (Jarrige, 1952;Anlaş, 2009;Assing, , 2013Anlaş et al., 2011;Örgel & Anlaş, 2016). This species is recorded from Afyonkarahisar, Aksaray, Denizli, Kayseri, Nevşehir ve Sivas provinces for the first time.
Distribution in the world: This species is widely distributed in Europe and also known from Middle Asia and Turkey (Schülke & Smetana, 2015).
Distribution in Turkey: A. turcicum was only known its type locality, Şarkikaraağaç, Isparta . Thus, this species is recorded for the first time outside the type locality and its distribution is found to be much wider than it was previously known. A. turcicum is recorded for the first time from Central Anatolia Region.
Remarks: For redescription of this species, see  Distribution in the world: This is an endemic species for Turkey (Schülke & Smetana, 2015).

DISSCUSSION
The genus Achenium Leach includes 54 species in the Palaearctic region . According to recent studies, the genus was represented by 11 species in the recorded species; A. humile from the Aegean Region and A. turcicum from Central Anatolia Region were recorded for the first time. A. turcicum was reported for the first time outside the type locality and its distribution was found to be much wider than previously known. A. debile was reported based on a old specimen from Konya province in central Anatolia region . But it was not recorded from central Anatolia in this study. Thus, this record is highly doubtful in central Anatolia.